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One the season’s most anticipated events is the upcoming four-way crossover between Arrow, The Flash, Supergirl, and DC’s Legends of Tomorrow. As the episodes in question begin production, we’ve now starting to learn more about the storyline.

One of the biggest questions has always been: what sort of villain could be so threatening that it would require the heroes of all four shows to fight it? After all, Supergirl doesn’t even take place in the same universe — it would require something truly overwhelming to draw her into the conflict.

Well, now we know. Today, Warner Bros. Television and The CW have finally revealed the answer: The Dominators. Here’s Arrow and Legends of Tomorrow executive producer Marc Guggenheim:

“This year, for our mega ‘Arrowverse’ crossover, we’re taking inspiration from a DC crossover from the late 1980s known as ‘Invasion!,’ which featured an alien race called the Dominators, who’d previously vexed the Legion of Super-Heroes. We’re using cutting-edge prosthetics and computer effects to achieve a feature film-quality look which is faithful to Invasion! artist Todd McFarlane’s interpretation of the characters.”

The Dominators

The Dominators first appeared in Adventure Comics #361 in October 1967. They are a highly technologically advanced race, capable of manipulating their own genes to enhance members of their species. The have a very distinctive appearance, with elongated hairless heads,yellow skin, and sharp, fearsome teeth. The size of a red circle on their foreheads indicates each Dominator’s place in their hierological society.

Invasion!

Invasion! was a three-part crossover event published in 1988 and 1989 by DC Comics. While the Dominators lead the titular invasion, they were part of a larger assembled coalition of races. One of those races were the Thanagarians — the same race that Hawkman and Hawkgirl come from. It’s possible Legends of Tomorrow could explore this angle.

Crossover Production is Underway

Arrow star Stephen Amell has been teasing the filming on the crossover on social media. Yesterday, he shared an image of himself with Supergirl star Melissa Benoist in costume:

Early this morning, Amell shared another photo after a long day of filming. The picture shows Amell posing with Flash star Grant Gustin and Legends stars Franz Drameh and Victor Garber:

We’ll, of course, have more details as they come in. Be sure to follow us on Facebook, Twitter, and Tumblr for all the latest news and updates.